2. PROLIFERATION CONTROL

This cellular division is strictly controlled by growth factors (GFs), growth inhibitors and other controlling molecules. Growth factors can act on cells in different phases of the cell cycle. Each cell responds to specific combinations of signalling molecules incl. GFs; different cells can respond differently to the same signal (e.g. growth factor). Examples of GFs are shown in the Table; at least two examples are required in final examination.

Growth factor Cells proliferating in response to GF
EGF
NGF
PDGF
FGF
IGF-1
IL-2
EPO
VEGF
HGF
epidermal GF
nerve GF
platelet-derived GF
fibroblast GF
insulin-like GF 1
interleukin-2
erythropoietin
vascular endothelial GF
hepatocyte GF
keratinocytes, trophoblast, neuroepithelial cells etc.
neuroblasts, neurite growth
connective tissue cells, neuroglia
many cell types
acts with other GFs
activovated T lymphocytes
precursors of erythrocytes
endothelial cells
hepatocytes

Growth factors can influence only those cells that possess specific receptors for the particular growth factor. The receptors for most growth factors are transmembrane Tyrosine-specific protein kinases. Through their activation, growth factors trigger cascades of intracellular signals and activate new genes. Some growth factors can also act as differentiation factors.

Although the proliferation is responsible for the growth of the embryo, there are many more mechanisms contributing to the growth of the body:

proliferation – hyperplastic growth,
enlargement of cell size – hypertrophic growth,
• production and accumulation of the extracellular matrix.

More pieces of information concerning “The Cell Cycle” can be found in the electronic course “Cytology II”.